It could have actually happened. The first time he auditioned for a Joss Whedon television show he was trying out for the part of Angel. According to Fillion (pictured on left) he didn't make it far enough through the auditions to actually meet the producers, so Whedon didn't see him then. Years later Whedon told him he wasn't even aware that he'd auditioned for the role (see clip at end).
It's clear that Whedon loves Fillion (as an actor, and I'm assuming also as a person). He's used him in multiple projects: in the final season of 'Buffy' (as Caleb), in 'Firefly' and 'Serenity', in 'Dr. Horrible's Sing-Along Blog', and in 'Much Ado About Nothing'.
Because it's clear that Whedon loves Fillion's talent (which I'm in no way disputing he has), I'm secretly glad that Fillion didn't make it to the final casting round for Angel.
Fillion is a brilliant actor, and he's definitely multi-faceted. He's also attractive or, as they say on 'Castle', "ruggedly handsome". But he doesn't have the dark, brooding handsomeness that helped David Boreanaz fill the role. Everyone uses "dark" and "brooding" as adjectives for Boreanaz (image on left). Seriously, just try Googling those words with his name.
If Fillion had been cast as Angel it would have impacted the entire tone of the Buffy and Angel relationship, as well as many of the show's central plots. I know that looks are in no way the most important factor of an actor's performance, but they do matter.
Considering the fact that Fillion and Boreanaz are on quite similar shows now ('Castle' and 'Bones'), I wonder if their careers paths would have still followed the same trajectory if Fillion had been cast as Angel instead of Boreanaz. Luckily, we'll never know.
I would've still loved to see Fillion as Angel, I always thought he (Angel) turned out a bit one dimensional. That same love Joss has for NF could've helped the part to get more multifaceted and challenging. On the other hand, we wouldn't have Mal and this role is the one NF was clearly born to play so all is good ))
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